 The longer breeding seasons in Southern Europe can also lead to reproductive capacity in worker bees, making "L. malachurum" in those regions not purely eusocial.Female workers in southern European colonies of "L. malachurum" consequently have significantly more developed ovaries than their counterparts in northern European colonies.Due to the relatively broad range of nesting habitats of the species, "L. malachurum" is subject to a myriad of climate-based selective pressures that cause a differential in behavior dependent upon location.