 However, in 1994 the ichnotaxon "Tyrannosauropus" were identified as large plant-eater dinosaur tracks, and when combined with analyses to distinguish three-toed carnivore and herbivore dinosaur footprints the large tracks attributed to "Tyrannosauropus" at Lark Quarry were interpreted as being produced by a large herbivore similar to "Muttaburrasaurus", rather than by a predatory theropod, and were suggested to belong to the ichnogenus "Amblydactylus" instead.Research by Anthony Romilio and colleagues casts doubt on the original interpretation.Analysis of the sediments indicates that they were deposited by a seasonal water course with water flowing at different depths and speeds at different times.