 It was this study that supported the split between "Nephilengys" and "Nephilingis".Although "Nephila" appears not to be monophyletic, the authors of the study did not suggest splitting the genus.The phylogeny suggests that male enforced monogamy, via plugging of the female copulatory ducts by males leaving behind their palpal bulbs, is ancestral to the nephilines, and was lost in "Nephila" and "Clitaetra".