 Born in Rome, graduated in agriculture and genetics, he entered the Superior Institute of Health in 1950, founding a department of Microbiological Genetics.He became professor of genetics at the University of Camerino, then at the University of Palermo in 1965, and finally moved to the University of Perugia in 1970, where he was emeritus professor and managed the Genetics Institute of the University from 1974.From 1970–1971 he presided over the "Associazione Genetica Italiana".