 The sediment containing the fossil pollen had accumulated in a lowland wetland environment with various vegetation units of mixed evergreen/deciduous broadleaved/conifer forests surrounding the wetland basin.Key relatives of the fossil taxa found with Distylium are presently confined to humid warm temperate environments, suggesting a subtropical climate during the middle Miocene in Austria.Distylium macrofossils of the Lower and Middle Miocene, are known from the lignite mines of the Kaltennortheim Formation in the Rhön Mountains, cental Germany, where it is associated with typical elements of the Mastixioid floras that attest to an optimal warm humid phase of the Miocene.