sentences:  It was first successfully breeding ex-situ for the Costa Rican frogs was at Manchester Museum's Vivarium  where an official conservation project for the Costa Rican population was established (Project Lemur Frog) was established in 1999.Work at Manchester Museum with the species has involved supporting them in Costa Rica, highlighting the species' plight to the public  Manchester University has also developed a global environmental education programme based on the species that extends to the countries of origin.The Atlanta Botanical Garden and El Valle, Panama have also bred the Panamanian form of the species in captivity and been highly successful.