sentences:  They are mainly native to freshwater habitats in South America, but there are also several species (in genera "Crossoloricaria", "Dasyloricaria", "Fonchiiichthys", "Rineloricaria", "Spatuloricaria", "Sturisoma" and "Sturisomatichthys") in Panama and a single ("Fonchiiichthys") in Costa Rica.Loricariinae was first described in 1831.Later, in 1979, many genera were described and Loricariinae was divided into four subfamilies