sentences:  He experimentally grew two species of "Bromus" along a moisture gradient and showed that, in monoculture,both species were able to inhabit the entire range.Nevertheless, in mixture, the two species dissociated and showed clear preference for either the dry or the moist end of the gradient
  
  In 1966 he moved to the University of Göttingen, where he established the Neuer Botanischer Garten der Universität Göttingen.He became emeritus in 1981.