sentences:  In 1872 he defended his doctoral thesis with a study on compound ascidians titled ""Recherches sur les ascidies composées ou synascidies"".From 1873 to 1882, he was "professeur suppléant" of natural history at the faculty of sciences in Lille, and in the meantime, was also affiliated with the Institut industriel du Nord.In 1874 he founded a biological station at Wimereux in order to familiarize his students to marine and terrestrial organisms.