sentences:  Witton suggested that istiodactylids would have been powerful fliers, due to an enlarged area for downstroke musculature attachments and well-developed pectoral and upper arm bones, and they probably spent much time in the air.Istiodactylids had wing-membranes connected to shortened bodies with short legs and long forelimbs, which may have created large wings with a high aspect ratio and low wing loadings.The wings of the istiodactylid "Nurhachius" have been compared to those of modern soaring birds (that fly with little flapping), and may have been ideal for low-energy soaring, which is necessary when searching for carrion.