sentences:  However, Inger in a 1948 paper found the specimens wanting of characteristics that would justify a generic separation from "Osteolaemus" and referred the specimens to "Osteolaemus osborni".In 1961, it was reduced to subspecies rank.A study of morphology published in 2007, and studies of DNA in 2009, 2013 and 2015 indicate that three distinctly different populations of "Osteolaemus" may merit full species recognition.