sentences:  It was collected by James Kitching and Regent "Lucas" Huma in sandstone that was deposited during the Hettangian stage of the Jurassic period, approximately 201 to 199 million years ago.The paratype material BP/1/5278 (originally assigned to "Syntarsus rhodesiensis") was discovered in 1981, also at the Elliot Formation in pinkish-maroon silty mudstone that was deposited in Hettangian sediments.Both the holotype and paratype specimen were housed in the fossil collection of the Evolutionary Studies Institute, part of the School of Geosciences of the University of the Witwatersrand, in Johannesburg, South Africa.