sentences:  Moroccan collections under the cork oak ("Quercus suber") that were initially regarded as "B. aereus", were described as a separate species—"Boletus mamorensis"—in 1978, on the basis of a rufous chestnut cap and a rooting stipe, or stem, with a reticulation often limited to the top (apex).However, molecular phylogenetic studies by Bryn Dentinger and colleagues in 2010, placed these collections very close to "B. aereus", suggesting they are more likely an ecological variant or phenotype, rather than a distinct species.More recent phylogenetic studies by M. Loizides and colleagues in 2019, have confirmed that "B. mamorensis" is a later synonym of "B. aereus", since collections identified as the two taxa could not be genetically separated and nested in the same clade.