Notocupes
Notocupes is an extinct genus of medium-sized archostematan beetles from the Triassic to Cretaceous. It is generally classified as a member of the family Ommatidae, but the presence of characters such as overlapping abdominal sternites indicate that the genus may have a closer affinity with the family Cupedidae. Notocupes is considered to be a junior synonym of Zygadenia by Kirejtshuk (2020), but other researchers suggest to reserve the genus Zygadenia as a form taxon for isolated elytra that probably belong to the genus Notocupes, while retaining Notocupes as a valid genus for complete body fossils. Notocupes has a flattened body, which may suggest that it occupied narrow habitats, such as living under bark. Some species had serrated/spined margins of the carapace, which may have served as a defense against predators, or served as camouflage to resemble bark. Most species of Notocupes were described from compression fossils. An additional three species were described from Cenomanian-aged Burmese amber, which are treated as a separate genus, Echinocups, by Kirejtshuk (2020). Li et al. (2023) consider Echinocups to be a junior synonym of Notocupes.

Triassic species

Ladinian (242 Ma to ~237 Ma)
 Tongchuan Formation, China
 N. shiluoensis

Carnian (237 Ma to 227 Ma)
Koldzat Formation, Kazakhstan
 N. oxypygus 
Madygen Formation, Kyrgyzstan
 N. laticella 
 N. rostratus  
 N. tenuis

Jurassic species

Hettangian (201.3 Ma to 199.3 Ma)
Dzhil Formation, Kyrgyzstan
 N. cellulosus  
 N. issykkulensis 
 N. kirghizicus 
 N. latus 
 N. sogutensis 
Guanyintan Formation, China
 N. homorus

Toarcian (182.7 Ma to 174.1 Ma)
Guanyintan Formation, China
 N. lini 
Sulyukta Formation, Kyrgyzstan
 N. crassus

Bajocian (170.3 Ma to 168.3 Ma)
Bakhar Formation, Mongolia
 N. brachycephalus 
 N. exiguus 
 N. longicollis

Callovian (166.1 Ma to 163.5 Ma)
Haifanggou Formation, China
 N. daohugouensis 
 N. dischides 
 N. jurassicus 
 N. pingi 
 N. robustus 
 N. spinosus 
Tyumen Formation, Russian
 N. patulus

Oxfordian (163.5 Ma to 157.3 Ma)
Karabastau Formation, Kazakhstan
 N. foersteri  
 N. lapidarius 
 N. nigromonticola 
 N. picturatus 
 N. pulcher

Tithonian (152.1 Ma To 145.0 Ma)
Shar Teeg, Mongolia
 N. brachycephalus  
 N. exiguus 
Solnhofen, Germany
 N. reticulatus ) 
 N. tripartitus

Cretaceous species

Hauterivian (132.9 Ma to 129.4 Ma)
Dabeigou Formation, China
 N. lentus

Barremian (129.4 Ma to 125.0 Ma)
Jianshangou Formation, China
 N. laetus 
La Huérguina Formation, Las Hoyas, Spain
 N. diazromerali  
 N. longicoxa   
 N. siniestri  
 N. viridis 
La Pedrera de Rúbies Formation, Las Hoyas, Spain 
 N. martinclosas  
 N. oculatus 
Ulan-Argalant Formation, Mongolia
 N. mongolicus

Aptian (125.0 to ~113.0 Ma) 
Argun Formation, Russia
 N. semen 
 N. sibiricus 
Dzun-Bain Formation, Mongolia
 N. dundulaensis  
 N. elegans 
Jehol Group, China
 N. validus  
Khasurty Formation, Russia 
 N. khasurtyiensis 
Laiyang Formation, China
 N. laiyangensis 
 N. ludongensis 
 N. tuanwangensis 
Lushangfen Formation, China
 N. trachylenus 
Shahai Formation, China
 N. kezuoensis 
Yixian Formation, China
 N. alienus 
 N. epicharis  
 N. eumeurus  
 N. minisculus 
 N. porrectus 
 N. protensus  
 N. psilatus  
 N. rudis 
 N. stabilis 
Zaza Formation, Russia
 N. caudatus 
 N. excellens 
 N. vitimensis

Albian (~113.0 to 100.5 Ma)
Jinju Formation, South Korea
 N. premeris

Cenomanian (100.5 to 93.9 Ma)
Burmese amber, Myanmar
 N. denticollis  (sometimes placed in separate genus Echinocups)
 N. neli  (sometimes placed in separate genus Echinocups)
 N. ohmkuhnlei  (sometimes placed in separate genus Echinocups)

Turonian (93.9 to 89.8 Ma)
Emanra Formation, Russia
 N. khetanensis  
Kzyl-Zhar Locality, Kazakhstan
 N. caducus

References

Ommatidae
Prehistoric beetle genera
Triassic insects
Jurassic insects
Cretaceous insects