 Complete the following phrase:  Glyptodonts first evolved during the Miocene in South America, which remained their center of species diversity. For example, an Early Miocene glyptodont with many primitive features, when compared to other species, Parapropalaehoplophorus septentrionalis, was discovered at a now-elevated site in Chile and described in 2007. After the Isthmus of Panama formed about three million years ago, the genus Glyptotherium spread north as part of the 