 To be clear, however, there can be indiscriminate sampling processes and discriminate sampling processes occurring in the same population, even with respect to the same trait. For example, in a study of over 900 populations, biologist Maxime Lamotte acknowledged that camouflage gave appropriately colored Cepaea nemoralis land snails a selective advantage in their respective environment while simultaneously maintaining that foundings of new populations are “of considerable importance because of the chance variations in the composition of the first colonizers” (Lamotte 1959: 80); the variations he refers to are variations in the colors of the snail colonizers (Millstein 2009). Moreover, since the Causal Process Account requires that variations be heritable, the non-heritable locations of organisms are simply irrelevant for the purposes of deciding on drift vs. selection; for example, the founders of a new population may all hail from the geographic edge of the original population, but they can still be an indiscriminate sample of the whole. Finally, historically, at least (as discussed above), small population size has always been associated with drift, but it was never the main phenomenon to be represented. Thus, the variant sampling accounts of drift should be evaluated in light of these considerations.