 Second, the description of the strategy picked out by the model is not simply a description of it as a series of behavioral dispositions (i.e. dispositions to lay different sex ratios under different conditions), but of the relationships between those dispositions and various ecological features that determine the fitness of those dispositions. For example, in the jewel wasp case, the presence of other female competitors, flexibility in brood size, the capacity of sons to mate multiple times, the impossibility of migration of sons between pupae and so on, are features of the environment that determine whether any potential egg laying strategy of the jewel wasp is maximally adaptive, and a well confirmed optimality model can show how. Consequently, contingent on obtaining the appropriate evidence that this is the correct model, the description of the strategy given by a successful model is (implicitly or explicitly) a functional description of the strategy—a description of how the behavior contributes to the fitness of the organism. For a more detailed discussion of the role of optimality modeling in biology, see the entry on  adaptationism.