 A prominent example is the issue of levels of selection. In this debate, the hierarchical organization of nature into levels is an important background assumption, as the aim is to find out at which level(s) of the biological hierarchy natural selection is taking place (Griesemer 2000; Okasha 2006). Although Darwin’s original account was focused on evolution at the level of organisms, arguably the conditions for natural selection can be formulated abstractly without referring to any specific kinds of entities, which allows for natural selection to operate at any level where the conditions are satisfied (Griesemer 2000; Lewontin 1970). Since the 1970s, the debate on levels of selection has kept on growing and extending to different areas, though no precise consensus has been reached. Positions range from the gene-centered view, where natural selection is taken to operate almost exclusively at the level of genes (e.g., Dawkins 1976; Williams 1966), to the pluralistic multilevel selection theory, which allows for natural selection to operate on any level of the biological hierarchy where we find the right kind of units (e.g., Sober &amp; Wilson 1998; Wilson &amp; Wilson 2008).