 One of the best arguments for studying heredity through the perspective afforded by multiple inheritance systems is that this perspective opens up questions about the evolutionary and developmental relations and interactions between the various inheritance systems that are characterized. Among these questions are questions about whether each system creates new targets of selection, about the ways in which inheritance systems may provide developmental scaffolding for other inheritance systems, about the regulatory role they may have in relation to one another, and about the evolution of inheritance systems. We now turn to these issues.