 Jablonka and Lamb (2005/2014) present a general account of biological evolution based on the multiple inheritance systems perspective. They argue that evolution can occur through any of the inheritance systems they identify (e.g., the behavioral) without necessarily involving genetic changes. This can happen through natural selection operating on non-genetic hereditary variations. Epigenetic changes are usually generated at a higher rate than genetic changes, often as a result of changes in environmental conditions, and the variation that is generated may have a higher chance of being beneficial than blind variation. This may allow rapid adaptation to changing conditions. These claims apply to behavioral inheritance and symbolic inheritance as well. Shea, Pen, &amp; Uller (2011) distinguish between the adaptation resulting from selection on epigenetic variations, which they term selection-based effects, and the adaptation resulting from induced response to the environment, which they term detection-based effects, and discuss their evolutionary and developmental consequences. Selection-based effects lead to adaptation via natural selection operating on reliably transmitted epigenetic variations and are analogous to the selection-based effects of genetic inheritance, though epigenetic variation may occur more rapidly and its frequency may increase due to environmental challenges. Detection-based effects, in contrast, are the result of directional variation and are a form of phenotypic plasticity.