  The individual elements of the matrix \(W_{ij}\) are the fitness consequences of response \(i\) in state \(j\).  So, for instance, \(W_{21}\) denotes the fitness consequences of \(R_2\) in \(S_1\). If we let \(W_{11}\) and \(W_{22}\) equal one and \(W_{12}\) and \(W_{21}\) equal zero, then there is a clear evolutionary advantage to performing \(R_1\) in \(S_1\) and \(R_2\) in \(S_2\).