 Maddison (1997) is also directly about how we ought to incorporate what we learn about the internal structure of lineages into our models of phylogenetic inference, and how we might build on the earlier work on coalescent models. Maddison is among the first to deeply appreciate how impactful the then-new work on genomics might be for phylogenetics, both in the ways that discordance will present serious challenges and opportunities for phylogenetic inference. This has helped spur new developments in this field of phylogenetics, (e.g., Maddison &amp; Knowles 2006; Degnan &amp; Rosenberg 2006, 2009; Baum 2007; Nakhleh 2013; Hahn &amp; Nakhleh 2016; Mallet, Besansky, &amp; Hahn 2016).