 Here \(H_T\) is the total heterozygosity of the population assessed, and \(\bar{H}_S\) is the heterozygosity of each subpopulation (group) of the relevant population, averaged across all the subpopulations. \(F_{ST}\) is bounded by 0 and 1, and is a measure of population structure, with higher \(F_{ST}\) values suggesting more structure, viz., more group differentiation. HWP appears implicitly in both \(H_T\) and \(\bar{H}_S\), which take heterozygosity (\(2pq\)) to be equal to the expected proportion of heterozygotes under HWP rather than the actual frequency of heterozygotes. \(H_T\) is computed by using the grand population average of \(p\) and \(q\), whereas calculating \(\bar{H}_S\) involves averaging across the expected heterozygosities of each subpopulation. If random mating occurs—and thus HWP applies—across the entire population without respecting subpopulation borders, then \(H_T\) and \(\bar{H}_S\) will be equal (i.e., \(p\) of the total population and of each individual subpopulation will be the same; likewise for \(q\)). If, instead, HWP applies only within subpopulations but not across the population as a whole, then \(\bar{H}_S\) will be smaller than \(H_T\), and \(F_{ST}\) will be positive (i.e., there will be “excess homozygosity” across subpopulations, which is known as the “Wahlund Principle” in population genetics). This is one way among many to deploy the population-genetic principle of HWP. Thus, the Lewontin-style diversity partitioning result that only roughly 5% of the total genetic variance is among races is equivalent to saying that \(F_{ST}\) across the big three continental populations in Lewontin’s three-level model is 0.05 (e.g., Barbujani et al. 1997). The basic philosophical tendency is to associate the diversity partitioning research project’s (approximately) 85%-10%-5% result with an anti-realist interpretation of biological race.