 As noted earlier, selection and drift are construed in logically distinct fashions in type recursions and the Price Equation. What is called is selection in type recursions is one sort of definite thing: it’s whatever it is that must be quantified by fitness variables. What is called selection in the Price Equation is another thing, and this will be determined by researchers’ understanding of the system, in particular how they generate expectations about fitness. Ultimately, the conflict between the two modeling approaches with respect to what counts as selection may be resolvable in at least a couple of ways. Perhaps one modeling approach is simply wrong about what selection is. Alternatively, something having to do with selection is arbitrary here. In particular, whether some or another form of randomness should be treated as selection or drift is a function of one’s choice of which modeling approach to use, where distinct approaches provide logically distinct understandings of “selection” and “drift”. Once one chooses to model the system using type recursions, one’s hand is forced: the temperature must be modeled as selection. Once one chooses to model the system using the Price Equation, one’s hand remains free: one may treat the changing temperature as relevant to expected fitness, and hence treat it as selection, or instead treat it as producing deviation from expectation, and thereby treat it as drift. According to this second way of resolving the conflict, the choice between the two modeling approaches is not dictated by nature, and is thus at least metaphysically (if not pragmatically) arbitrary.