 Importantly, Millikan’s second core claim is a fully general thesis, and is thus intended to apply to systems of representations that are more complex than the system of beaver splash signals. Take one of Millikan’s favorite examples, the waggle dance of the honey bee. Different waggle dances do not only vary with respect to the time and place of their performance, as beaver splashes do, but also with respect to the length and direction of the ‘waggle run’. Furthermore, variations of the latter kind make a significant difference to the responses of the consumer bees: the length and direction of their subsequent flights in search of food Normally depends on the length and direction of the waggle run. Hence, we get a more complex mapping rule that takes us from dances with different waggle run lengths and directions (performed in location l at time t) to states of the form nectar source located in direction r at distance d (relative to location l, at  time t).[11]  This is the rule that specifies the correspondence relations that must obtain between dances and nectar positions for the consumer bees’ behavior to fulfill its biological functions (in the Normal way).