 In the light of these scientific findings, contemporary science and religion authors have reconsidered the questions of human uniqueness, imago Dei, the Incarnation, and the historicity of original sin. Some authors have attempted to reinterpret human uniqueness as a number of species-specific cognitive and behavioral adaptations. For example, van Huyssteen (2006) considers the ability of humans to engage in cultural and symbolic behavior, which became prevalent in the Upper Paleolithic, as a key feature of uniquely human behavior. Other theologians have opted to broaden the notion of imago Dei. Given what we know about the capacities for morality and reason in non-human animals, Celia Deane-Drummond (2012) and Oliver Putz (2009) reject an ontological distinction between humans and non-human animals, and argue for a reconceptualization of the imago Dei to include at least some nonhuman animals. Joshua Moritz (2011) raises the question of whether extinct hominin species, such as Homo neanderthalensis and Homo floresiensis, which co-existed with Homo sapiens for some part of prehistory, partook in the divine image.