 Godfrey-Smith’s approach to evolutionary individuals is intended to contrast with the earlier replicator-based views developed by Richard Dawkins and others. Replicator views have been central to discussions of the levels and units of selection (Godfrey-Smith 2015; Sober &amp; Wilson 1994, 1998; Okasha 2006), and derivatively so to views of biological individuals. Here genes are paradigmatic replicators, being housed in interactors, such as organisms, and it is the survival of these replicators that matters in evolution. The replicator framework emphasizes the importance for natural selection of high-fidelity copying across generations. By contrast, what matters for natural selection in Godfrey-Smith’s view is the establishment of parent-offspring lineages that feature heritability, as well as the variety of forms that reproduction can take in the establishment and stabilization of those lineages. 