 The continued scholarly interest in the origins of Darwin’s theory reflects not only the concerns of historians of science, but also those who see a continued relevance of Darwin’s own writings as sources of creative reflection for contemporary work in evolutionary biology (Gayon 2003). This phenomenon, however, presents difficulties for the historical understanding of Darwinism. Particularly within anglophone philosophy of biology, the emphasis on the lines of the development of Darwin’s evolutionary theory that have led to the consensus position achieved in the so-named “Synthetic” theory of evolution of the 1930s (Smocovitis 1996; Mayr &amp; Provine [ed.] 1980; Provine 1971), has tended to obscure the complex history of Darwin’s own theoretical reflections and the history of Darwinian theory since 1859 (Depew &amp; Weber 1995; Bowler 1983).