 One may understandably think that the early genic selectionists were interested in the replicator question because of the claims that the unit of selection ought to be the replicator. This would be a mistake. Rather, the primary interest is in a specific ontological issue about benefit (Dawkins 1976, 1982a,b). This amounts to asking a special version of the beneficiary question, and the answer to that question dictates the answers to the other three questions flying under the rubric of the “units of selection”.