 Lorenz responded to his critics with a revised account of the innate/acquired distinction (Lorenz 1965; see also Browne 2005). No trait is innate in itself, but “certain parts of the information which underly the adaptedness of the whole, and which can be ascertained by the deprivation experiment, are indeed innate” (Lorenz 1965, 40). A trait is innate insofar as its development is guided by ‘inherited information’ rather than ‘environmental information.’ Lorenz defined ‘information’ in terms of adaptation. An adaptive trait ‘fits’ its environment and hence can be said to contain information about that environment just as a key can be said to contain information about the lock it will open. The question is where that information comes from – how does the organism ‘know’ about its environment? If a woman has calluses on her palms rather than on the backs of her hands, this represents information about where her skin gets rubbed most. If she were born with calluses on her palms, as ostriches are born with calluses that match pressure points on their legs, this would imply advance ‘knowledge’ of where her skin is going to be rubbed most. Such anticipatory information, Lorenz argued, must be in the genome. It is in the genome as a result of natural selection, which can be seen as a form of trial-and-error learning.