 To the extent that active germ-line replicators benefit from the survival of the group of individuals in which they sit, over and above the [effects of individual traits and altruism], we may expect to see adaptations for the preservation of the group. But all these adaptations will exist, fundamentally, through differential replicator survival. The basic beneficiary of any adaptation is the active germ-line replicator (Dawkins 1982b: 85).