 Another, related issue arose concerning the character of the traits that were said to be subject to random drift. Wright’s early writings, in particular, emphasized the nonadaptive character of the traits undergoing drift and produced by drift (as can be see in Skipper’s characterization of Wright’s SBT above). But a number of potential confusions arise here (Provine 1986; Millstein 2009): First, “nonadaptive” does not necessary mean “maladaptive”; it can also mean “neutral”. Second, “nonadaptive” can refer either to the lack of adaptive differences between organisms or species, or it can refer to the lack of adaptation to the environment (but authors are not always clear about which meaning is intended). So, one’s views on the prevalence of drift can be—and historically often have been—tied to one’s beliefs about the prevalence of adaptive traits (in either sense) in populations. Strictly speaking, however, there is no necessary connection between nonadaptive traits and drift; there can be indiscriminate sampling on organisms that are adapted to their environment and which differ in fitness, and the outcomes of that indiscriminate sampling can pull the population in an adaptive direction or a nonadaptive one. Thus, the proper contrast between selection and drift does not map onto “adaptive” and “nonadaptive”, but rather to “adaptive” and “possibly nonadaptive” (but also “possibly adaptive”).