 Evolutionary game theory has been used to explain a number of aspects of human behavior. A small sampling of topics which have been analysed from the evolutionary perspective include: altruism (Fletcher and Zwick, 2007; Gintis et al., 2003; Sánchez and Cuesta, 2005; Trivers, 1971), behavior in public goods game (Clemens and Riechmann, 2006; Hauert, 2006; Hauert et al., 2002, 2006; Huberman and Glance, 1995), empathy (Page and Nowak, 2002; Fishman, 2006), human culture (Enquist and Ghirlanda, 2007; Enquist et al., 2008), moral behaviour (Alexander, 2007; Boehm, 1982; Harms and Skyrms, 2008; Skyrms 1996, 2004), private property (Gintis, 2007), signaling systems and other proto-linguistic behaviour (Barrett, 2007; Hausken and Hirshleirfer, 2008; Hurd, 1995; Jäger, 2008; Nowak et al., 1999; Pawlowitsch, 2007, 2008; Skyrms, 2010; Zollman, 2005), social learning (Kameda and Nakanishi, 2003; Nakahashi, 2007; Rogers, 1988; Wakano and Aoki, 2006; Wakano et al., 2004), and social norms (Axelrod, 1986; Bicchieri, 2006; Binmore and Samuelson, 1994; Chalub et al., 2006; Kendal et al., 2006; Ostrum, 2000).