 Armed with these characterizations, Von Eckardt Klein argues that inference to a functional-localization hypothesis proceeds in two steps. First, a functional deficit in a patient is hypothesized based on the abnormal behavior the patient exhibits. Second, localization of function in normal brains is inferred on the basis of the functional deficit hypothesis plus the evidence about the site of brain damage. The structurally-adequate functional analysis of the capacity connects the pathological behavior to the hypothesized functional deficit. This connection suggests four adequacy conditions on a functional deficit hypothesis. First, the pathological behavior P (e.g., the speech deficits characteristic of Broca’s aphasia) must result from failing to exercise some complex capacity C (human speech production). Second, there must be a structurally-adequate functional analysis of how people exercise capacity C that involves some constituent capacity ci (formulating motor commands to produce the appropriate sounds). Third, the operation of the steps described by the structurally-adequate functional analysis minus the operation of the component performing ci (Broca’s area) must result in pathological behavior P. Fourth, there must not be a better available explanation for why the patient does P. Arguments to a functional deficit hypothesis on the basis of pathological behavior is thus an instance of argument to the best available explanation. When postulating a deficit in a normal functional component provides the best available explanation of the pathological data, we are justified in drawing the inference.