 Some degree of reintegration of what is unobservable in the classical use of quantitative genetic models has occurred through the technique of mapping quantitative trait loci (QTL)—regions of the genome containing genetic factors associated with variation in a continuously variable trait. Yet QTL mapping has had most success in animal and plant varieties that can be replicated and raised in controlled conditions; reliable QTL results for human populations are few (Majumder &amp; Ghosh 2005; but see the Wellcome Trust Case Control Consortium 2007). Indeed, reintegration of the complexities of development in its ecological context, which are collapsed into the parameters of the models, remains scarcely developed in quantitative genetics (as evoked for the human case by Turkheimer 2004).