 While large amounts of work in cultural evolution have focused on the human species, there is also a growing body of work assessing the implications of learning for adaptation and speciation in many other species including chimpanzees (Whiten et al. 1999), whales (Rendell and Whitehead 2001), fish and birds among many others (Laland and Hoppitt 2003). Moreover, this work on non-human species also helps to refine and to answer a series of questions about why humans, compared with other species, seem so conspicuously good at building, maintaining and refining collective storehouses of adaptive cultural capital (Henrich 2015, Laland 2016).