 Non-genetic inheritance can have an effect on the ecological conditions an organism faces by affecting or determining behavior and activities, thus increasing or dampening selection. The result is a feedback loop between actions of the organism and selection that leads to what Conrad Waddington referred to as the cybernetic nature of evolution (Waddington 1961). Behavioral and symbolic inheritance, in particular, can reinforce this process. By thus affecting the selective challenges faced by the organism, the evolutionary feedback loop can turn short-term hereditary effects, that would not survive many generations, into long term evolutionary change.  Section 4.4  further discusses the ramifications of this phenomenon.