 Let’s first consider the appeal of including molecular data in phylogenetic analyses—perhaps even to the exclusion of morphological characters (e.g., Scotland, Olmstead, &amp; Bennett 2003 though see Wiens 2004). One of the most important aspects of the molecular revolution for phylogenetics was the sheer amount of new data available to systematists, including for groups such as bacteria, where fossil data were scarce or severely lacking. Yet it was not merely the scale and availability of molecular data that held such appeal. As it became easier to collect, code, and collate these data, biologists argued that molecular phylogenetics provided an independent source of evidence from morphological studies that bolstered evidence for common ancestry and patterns of phylogeny (Zuckerkandl &amp; Pauling 1965). In contrast, more traditional systematists such as Theodosius Dobzhanksy, Ernst Mayr and George Gaylord Simpson were skeptical of molecular evolution studies quite generally, and, in particular, the idea that they were in some way superior or could replace morphological studies (Dietrich 1998).