 In contrast to the preceding approach, we can separate the interactor and manifestor-of-adaptation questions in our group selection models (Wright 1980; see Lewontin 1978; Gould &amp; Lewontin 1979). This is done by distinguishing between what is called “intergroup selection,” that is, interdemic selection in the shifting balance process, and “group selection for group advantage” (Wright 1980: 840; see Wright 1929, 1931). The term “altruist” originally denoted, in genetics, a phenotype “that contributes to group advantage at the expense of disadvantage to itself” (1980: 840; Haldane 1932). This earlier debate is connected to the main group selection debate in the 1960s, in which the group selectionists asserted the evolutionary importance of “group selection for group advantage” (Wright 1980). The argument is that the primary kin selection model is “very different” from “group selection for the uniform advantage of a group”(1980: 841; like Arnold &amp; Fristrup 1982; Damuth &amp; Heisler 1988; Heisler &amp; Damuth 1987). There are excellent summaries of the empirical and theoretical discoveries enabled by “intergroup” selection models (Goodnight &amp; Stevens 1997; Wade 2016).