   Consider all the possible daughter lineages or lines of descent from some individual, \(o\). Call each of the possible lineages \(\omega\) and the set of them \(\Omega\). For each lineage \(\omega_i\) the probability that \(o\)’s actual daughter population is \(\omega_i\) will be \(Pr(\omega_i)\). \(F\), the fitness of \(o\), will not be given by weighted sums of possible daughter populations at a specific generation in the future, but rather by this value over the long run, as the number of generations increases without limit.  The definition of individual fitness, \(F\), is then given by the following equation (Pence and Ramsey 2013, 862; equation (4)): 