 Note that the “interactor question” does not involve attributing adaptations or benefits to the interactors, or indeed, to any candidate unit of selection. Interaction at a particular level involves only the presence of a trait at that level with a special relation to genic or genotypic expected success that is not reducible to interactions at a lower level. A claim about interaction indicates only that there is an evolutionarily significant event occurring at the level in question; it says nothing about the existence of adaptations at that level. As we shall see, the most common error made in interpreting many of the interactor-based approaches is that the presence of an interactor at a level is taken to imply that the interactor is also a manifestor of an adaptation at that level.