 The holobiont—the combination of the host and its microbiota—functions as a unique biological entity anatomically, metabolically, immunologically, and developmentally (Gilbert, Rosenberg, &amp; Zilber-Rosenberg forthcoming; Gilbert 2011). Similarly, a holobiont is seen as an “integrated community of species, [which] becomes a unit of natural selection” (Gilbert, Sapp, &amp; Tauber 2012: 334). That is, in essence, theorists claim that the holobiont can function as an interactor since it has features that bind it together as a functional whole in such a way that it can interact in a natural selection process. So what ties the different species together to produce an interactor? According to pioneering philosophical thought on holobionts and symbionts, it is the community’s common evolutionary fate, its being a “functioning whole,” that characterizes it as an evolutionary interactor, “objects between which natural selection selects” (Dupré 2012: 160; see also Dupré &amp; O’Malley 2013; Zilber-Rosenberg &amp; Rosenberg 2008). This community can also be described as a “team” of consortia undergoing selection (Gilbert et al. forthcoming). Others describe them as “collaborators” or “polygenomic consortia”, which has the advantage of encompassing both competition and cooperation within the holobiont (Dupré &amp; O’Malley 2013: 314; Lloyd forthcoming; see also Huttegger &amp; Smead 2011 on stag hunt game-theoretic results regarding the range of collaboration).