 Grush’s (2001, 2004) interest in temporal representation is part of his broader systematic project of addressing a semantic problem for computational neuroscience, namely: how do we demarcate study of the brain as an information processor from the study of any other complex causal process? This question leads back into the familiar territory of psychosemantics (see  section 3 above),  but now the starting point is internal to the practices of computational neuroscience. The semantic problem is thereby rendered an issue in philosophy of neuroscience, insofar as it asks: what does (or should) “computation” mean in computational neuroscience?