 We live now in the postgenomic era. Just what “postgenomic” means, though, is a point of contention. On the one hand, many life sciences have embraced genomics, bringing DNA sequencing methods and an attention to genes into their domain—conservation genomics, cancer genomics, behavioral genomics, immunological genomics, bacterial genomics, and marine genomics, to name just a few (entries:  biodiversity;  philosophy of immunology;  conservation biology).  On the other hand, it’s become abundantly clear that there is a great deal of complexity between a sequence of DNA and a trait, and that there are far more variables affecting that trait than just the order of As, Cs, Ts, and Gs. This has spawned the proliferation of other “omic” disciplines that attempt to catalogue and functionally understand the great variety of molecular and cellular entities and processes that contribute to the structure and function of organisms—proteomics, transcriptomics, metabolomics, lipidomics, and glycomics, to name just a few. What’s more, genome-wide association studies, which scour entire genomes for regions that are associated with traits, have revealed that in most case, tens, hundreds, or thousands of genomic regions are implicated in even fairly simple human traits like height. This tension—between an urge to trace things down to genes and a realization once you get there that there is far more to the story than genes—is at the heart of many philosophical reflections on genetics (Richardson and Stevens 2015; Reardon 2017; entries:  genomics and postgenomics;  philosophy of systems and synthetic biology;  heritability).  