 But these mechanism descriptions are often embedded in different developmental contexts (at different times in ontogeny) with distinct compositional relations (within and between species). The reciprocal interplay between genetic and physical difference makers is not maintained precisely because these compositional differences alter relationships of physical causation (fluid flow, tension, etc.; see  Section 1.3).  Developmental biologists have been able to generalize relationships of genetic causation (in terms of genetic mechanisms; see  Section 1.3)  across species quite widely, but the attempt to combine these with physical causation has necessitated narrowing the scope of the causal claims.