 Phylogenetic inference is, generally, best categorized as a problem of epistemology. It is, after all, a problem of inference, albeit specialized to how we justify or ground our beliefs about phylogeny. An epistemologist who wonders how we can have knowledge of the past can hardly do better than to look to phylogenetics (Sober 1988b). And as we have seen, philosophy of statistics and of inductive inference more generally looms large. But moving beyond epistemology, any philosopher interested in how science works can learn from phylogenetics. Here we will look at the cases of metaphysics and the general philosophy of science.